Most plant cells contain one or more membrane-bound vesicles called vacuoles. Within the vacuole is the cell sap, a water solution of salts and sugars kept at high concentration by the active transport of ions through permeases in the vacuole membrane. Proton pumps also maintain high concentrations of protons in the vacuole interior. These high concentrations cause the entry, via osmosis, of water into the vacuole, which in turn expands the vacuole and generates a hydrostatic pressure, called turgor, that presses the cell membrane against the cell wall. Turgor is the cause of rigidity in living plant tissue.
In the mature plant cell, as much as 90 percent of cell volume may be taken up by a single vacuole; immature cells typically contain several smaller vacuoles.
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