Aspects of this topic are discussed in the following places at Britannica.
...scales of either the ganoid or the cosmoid type. Cosmoid scales have a hard, enamel-like outer layer, an inner layer of cosmine (a form of dentine), and then a layer of vascular bone (isopedine). In ganoid scales the hard outer layer is different chemically and is called ganoin. Under this is a cosmine-like layer and then a vascular bony layer. The thin, translucent bony scales of modern fishes,...
...Fish scales are formed of bone from the deeper, or dermal, skin layer. The elasmobranchs (e.g., sharks) have placoid scales; these are bony, spiny projections with an enamel-like covering. Ganoid scales, which are found on such fishes as gars and the bowfin, are similar to placoid scales but are covered with a peculiar enamel-like substance called ganoin. It is thought that true teeth...
in integument: Fishes )...1). They may be divided into several types on the basis of composition and structure. Cosmoid scales, characteristic of extinct lungfishes and not found in any fishes today, are similar to the ganoid scales of living species. Placoid scales (or denticles) are spiny, toothlike projections seen only in cartilaginous fishes. Ganoid scales, sometimes considered a modification of the placoid...
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